Galápagos Islands – In Darwin’s Footsteps (Eastern Route)

9-day all-inclusive Galápagos Expedition Cruise with round-trip flights from Quito

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Nazca Booby birds seen on the Española island, Galapagos. Photo: Dennis Ballesteros
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Visit the UNESCO-protected Galápagos Islands, discovering their unique wildlife, flora and fauna.

Join us on a journey of ecological discovery as we travel in Charles Darwin's footsteps, seeing the natural wonders that so inspired him.

Explore the islands of San Cristóbal Sante Fe, Santa Cruz and Española, and spend two nights in Quito, one of South America's most charming cities.

Setting out from beautiful Quito 

Quito is one of South America’s most alluring capitals and a World Heritage City of graceful Spanish-era architecture, packed with cosy restaurants and artisanal shops. With two overnights here, you’ll have time to taste delicious Ecuadorian specialties and stroll through colorful markets brimming with local handicrafts. On the second day, you’ll visit Quito´s historic centre on a guided tour.  

Awe inspiring Galápagos Islands 

After exploring Quito´s historic centre and visiting the Equator, we’ll head to the Galápagos Islands for an unforgettable sea adventure. We’ll be on the lookout for iconic animals such as giant tortoises, marine iguanas and Blue-footed Boobies. On this active cruise, and with an onboard Expedition Team of local guides, you'll get closer to the many natural wonders in our small expedition boats, kayaks and glass-bottom boat.

9 days
50 departures between Feb 2026 and Mar 2028
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Key Expedition Details

9 days
50 departures between Feb 2026 and Mar 2028
International flights included
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From
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Galápagos Islands – In Darwin’s Footsteps (Eastern Route)

Itinerary

Your adventure at a glance. Daily highlights, included activities, and optional extras to make it unforgettable.

Begin your adventure in the vibrant city of Quito

Your journey begins on the Equator in the lush Andean foothills. At 9,350 feet, Quito is the second-highest capital city in the world, and its main draw is the exquisite Old Town which was founded on an ancient Inca settlement. A UNESCO World Heritage site, it’s packed with well-preserved colonial architecture as evidenced in colourful plazas, magnificent churches and handsome monuments.

Settle into your hotel and enjoy an overnight stay.

If you have time, why not come a few days earlier and join one of our pre-voyage options.

Straddle the Equator in the Middle of the World City

Today, enjoy a guided tour of Quito’s remarkable Old Town, one of the largest and best-preserved colonial-era quarters in South America. Wander through cobblestone streets as you admire the architecture and sample delicious chocolate made from the finest Ecuadorian cacao along the way.

Next, you’ll head to the iconic ‘Middle of the World City’, located around 16 miles north of Quito. After lunch, you’ll visit the famous monument that delineates the imaginary line that is the Equator. Have your camera ready to capture the moment you stand with one foot in each hemisphere.

Get ready to explore the spectacular Galápagos Islands

After breakfast, a short flight will take you to Baltra Island, a part of the spectacular Galápagos Archipelago. A transfer will take you straight to the port, where your comfortable expedition ship MS Santa Cruz II awaits you.

Get your first glimpse of the islands’ incredible wildlife on Mosquera Islet, home to one of the largest sea lion colonies in the archipelago. We’ll take our small panga boats ashore for a closer look.

As you stroll across the flat, sandy dunes, look out for blue-footed boobies and Sally Lightfoot crabs, and learn about the marine and terrestrial ecosystems from your naturalist guides.

Meet the famous giant tortoises

Your first stop today is the Cerro Colorado Tortoise Reserve, a breeding center for the endangered giant tortoise. It’s also home to other endemic species, including the San Cristóbal mockingbird and the lava lizard.

The giant tortoise is probably the most iconic of all Galápagos wildlife, with a lifespan of over 100 years. They thrive here, ambling around forests similar to their natural habitat.

Up to August 2024 we’ll call at Cerro Brujo, on the island’s west coast. With its white coral beaches, you’ll literally be walking in Darwin’s footsteps as the naturalist visited here in 1835. There are plenty of sea and shore birds to look out for here, as well as sea lions.  

After August 2024* we’ll instead likely explore Punta Pitt in our small panga boats. This is the only site in the Galápagos Islands where you might see three species of boobies. Enjoy a nature walk, explore the coastline by boat, or snorkel and swim in the crystal-clear water.    *Note – this is dependent on local authority restrictions.

Spot wildlife against idyllic backdrops

We start the day with a scenic nature walk on the blissful beaches of Santa Fe, home to the endemic Barrington land iguana and a colony of sea lions.

You can swim or snorkel from our small panga boats, take a kayak for a paddle around the island, or join a trip in our glass-bottomed boat to observe the marine life below. After a delicious lunch prepared by our onboard chefs, we head to South Plaza Island.

Keep an eye out for frigatebirds, swallow-tailed gulls, and Galápagos shearwaters gliding on the breeze. Here, among a dazzling array of flora, land iguanas laze by the shore while marine iguanas slip in and out of the water.

Learn about local conservation efforts on Santa Cruz Island

Today, we pay a visit to the Charles Darwin Research Station in Puerto Ayora.

At this giant tortoise breeding center, you can see recently hatched tortoises and Darwin’s famous finches – the birds he used as a basis for his theory of evolution by natural selection. There’s also an impressive forest of prickly pear cacti that is home to many land birds.

Enjoy a cooler climate over lunch in the lush highlands, where giant tortoises shuffle about munching on grass and wallowing in pools.

You’ll also have time for another activity, such as mountain biking or visiting a sugar plantation.

Lava lizards and incredible birdlife

Your Galápagos Islands cruise reaches Española, the archipelago’s southernmost island. Birdlife here is truly extraordinary. On cruises from April through December, look for Nazca boobies, blue-footed boobies, swallow-tailed gulls, and the mighty waved albatross.

This is also a great chance to spot red Española lava lizards and marine iguanas, as well as see El Soplador, the famous blowhole that blasts water 75 feet into the air.

After lunch on board the ship, we’ll head to the picturesque coral beach of Gardner Bay and nearby Osborn Islet to seek out sea lions, mockingbirds, and finches. If you’re feeling active, you can go snorkeling or kayaking.

Green turtles, sharks, and rare bird colonies

The abundant wildlife-watching opportunities continue at Eden Islet, where we hope to see Galápagos green turtles, rays, and friendly reef sharks. On shore, look for distinctive frigatebirds with their rubbery, red throats.

You can swim or snorkel to explore the reefs. You can also see the reefs from our glass-bottomed boat or by kayak. After lunch, we’ll sail to North Seymour.

The island is a great site for spotting colonies of blue-footed boobies and swallow-tailed gulls. Sea lions and marine iguanas also congregate here, attracted by the bountiful sea life in its waters.

From the Galápagos Islands back to Quito

Your expedition cruise draws to a close as you disembark at Baltra Island. A transfer will take you to the airport in Baltra for your flight to Quito – or if you prefer, to Guayaquil – to to connect to your international flight. To explore more of Ecuador, why not extend your stay and join one of our post-voyage options.

As you leave this fascinating region, you'll take with you incredible memories of amazing wildlife and wonderful experiences few have the privilege of seeing.

What else is included?

Hotel

Two nights at a hotel in Quito before the expedition cruise, including breakfast and dinner on Day 2

Flights

Flights in economy class between Quito and Baltra, including all taxes

Excursion

Full-day excursion: Historic Quito & Middle of the World, including lunch, before the expedition cruise 

What's not included?

  • International flights

  • Additional overnight(s) if required due to international flight timings 

  • Travel protection

  • Baggage handling

Optional extras

Enhance your journey with optional extras. Some activities are only bookable onboard and can’t be reserved online or by phone. Subject to availability.

Our Galápagos Islands ship

Get a taste of what life is like on board the ships that sail on this itinerary

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Offers

Enjoy limited-time savings on selected sailings. Combinable on select offers.

$200 off your first booking

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